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Barna Páll-Gergely, András Hunyadi, Jonathan Ablett, Hào Văn Lương, Fred Naggs, Takahiro Asami
Zookeys
Figure 39.Locations mentioned in literature on plectopylid taxonomy. 1 Muong-Bo 2 Phony-Tho 3 Trinh-Thuong 4 Muong-Hum 5 Nat-Son (Nhat Son) 6 Muong-Kong 7 Long-Ping 8 Pac-Kha 9 Ha-Giang 10 Tinh-Tuc 11 Cao-Bang 12 Déo-Ma-Phuc 13 Quang-Huyen 14 Ha-Lang 15 Cho-Ra 16 Bac-Khan 17 Nac-Ri 18 That-Khé 19 Cho-Moi 20 Lang-Son 21 Mansongebirge 22 Than-Moi 23 Bac-Ninh 24 Dong-Trieu 25 Bah-Mun 26 Kebao 27 Baie d’Along. The locations of “Col de Nuages” (Clouds Pass) could not be located. It is probably situated on Lao Kay Province, close to Muong-Hum.
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Barna Páll-Gergely, András Hunyadi, Jonathan Ablett, Hào Văn Lương, Fred Naggs, Takahiro Asami
Zookeys
Figure 3.Shells of Vietnamese Gudeodiscus species. A Gudeodiscus (Gudeodiscus) fischeri (Gude, 1901), Vn10-120, coll. PGB B Gudeodiscus (Gudeodiscus) fischeri, MNHN 24579 (holotype of Plectopylis fischeri) C Gudeodiscus (Gudeodiscus) fischeri, MNHN 24587 (holotype of Plectopylis tenuis) D Gudeodiscus (Gudeodiscus?) infralevis (Gude, 1908), MNHN 24604 (holotype of Plectopylis infralevis) E Gudeodiscus (Gudeodiscus?) infralevis, MNHN 24585 (holotype of Plectopylis soror). Photos: B. Páll-Gergely (A) and T. Deli (B–E). Scale represents 10 mm.
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Barna Páll-Gergely, András Hunyadi, Jonathan Ablett, Hào Văn Lương, Fred Naggs, Takahiro Asami
Zookeys
Figure 9.Shells (A–B) and apertural views (C–R) of Vietnamese Gudeodiscus, Sicradiscus and Halongella gen. n. species. A Gudeodiscus (Gudeodiscus?) suprafilaris (Gude, 1908), MNHN 24586 (holotype?) B Gudeodiscus (Gudeodiscus?) suprafilaris, 2011/81 C Gudeodiscus (Gudeodiscus) phlyarius (Mabille, 1887), Vn11-156 D Gudeodiscus (Gudeodiscus) phlyarius (Mabille, 1887) (typical “anterides/gouldingi”), MNHN-IM-2012-2164 E Gudeodiscus (Gudeodiscus) messageri messageri (Gude, 1909), MNHN-IM-2012-2215 F Gudeodiscus (Gudeodiscus?) hemmeni Páll-Gergely & Hunyadi, sp. n., Vn10-103A G Gudeodiscus (Gudeodiscus?) anceyi (Gude, 1901), GS22 H Sicradiscus mansuyi (Gude, 1908), 20081116C I Gudeodiscus (Gudeodiscus) giardi giardi (Fischer, 1898), 2011/81 J Gudeodiscus (Gudeodiscus) villedaryi (Ancey, 1888), Vn11-151 K–L Gudeodiscus (Gudeodiscus) dautzenbergi (Gude, 1901), Vn10-44 M–N Halongella schlumbergeri (Morlet, 1886), MAA3 O Halongella fruhstorferi (Möllendorff, 1901), Vn11-171 P Gudeodiscus (Gudeodiscus) fischeri (Gude, 1901), Vn10-120 Q Gudeodiscus (Gudeodiscus) fischeri (Gude, 1901), 20090515C R Gudeodiscus (Gudeodiscus?) suprafilaris (Gude, 1908), 2011/81. All photos by B. Páll-Gergely except for Figure 9A (T. Deli). Scale represents 10 mm and refers to A and B.
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Barna Páll-Gergely, András Hunyadi, Jonathan Ablett, Hào Văn Lương, Fred Naggs, Takahiro Asami
Zookeys
Figure 15.Parietal (A, C, E, F, H, J, L, N, O, Q) and palatal (B, D, G, I, K, M, P, R) plicae and lamellae of Gudeodiscus species. A–D Gudeodiscus (Gudeodiscus) infralevis (Gude, 1908) A–B holotype of Plectopylis infralevis (after Gude 1908) C–D Holotype of Plectopylis soror (after Gude 1908) E–G Gudeodiscus (Gudeodiscus?) cyrtochilus (Gude, 1909) E–F MNHN-IM-2012-2251 (two different specimens) G 2012/47 H–N Gudeodiscus (Gudeodiscus) fischeri (Gude, 1901) H–I Plectopylis fischeri (after Gude 1901a) J–K 20090515C L–M Vn10-120 N MNHN-IM-2012-2241 O–R Gudeodiscus (Gudeodiscus) fischeri (Gude, 1901) (identical with the holotype of tenuis) O–P after Gude (1901a) Q–R Vn10-28A (two different specimens). Inner views: B, D, I, M, P; Outer views: G, K, R.
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Barna Páll-Gergely, András Hunyadi, Jonathan Ablett, Hào Văn Lương, Fred Naggs, Takahiro Asami
Zookeys
Figure 17.Reproductive anatomy of Gudeodiscus (Gudeodiscus) fischeri (Gude, 1901). Locality information: Tuyên Quang Province, near Ton Hông, road #185 from Tuyên Quang to Vĩnh Lộc (formerly Chiêm Hóa) (NE of Tuyên Quang), leg. Hemmen, Ch. & J., 19.03.2011. Scale represents 5 mm.
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Barna Páll-Gergely, András Hunyadi, Jonathan Ablett, Hào Văn Lương, Fred Naggs, Takahiro Asami
Zookeys
Figure 18.Reproductive anatomy of Gudeodiscus (Gudeodiscus) fischeri (Gude, 1901) (typical tenuis specimen). Locality information: Bắc Kạn Province, Ba Bể Nat. Park, Hang Thẳm Kit 2 km from the look-out tower, 335 m, 22°24.686'N, 105°37.710'E, leg. Hunyadi, A., 19.11.2011. Scale represents 5 mm.
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Barna Páll-Gergely, András Hunyadi, Jonathan Ablett, Hào Văn Lương, Fred Naggs, Takahiro Asami
Zookeys
Figure 28.Inner walls of the penis of Gudeodiscus Páll-Gergely, 2013 species. A Gudeodiscus (Gudeodiscus) phlyarius (Mabille, 1887) (typical fallax specimen, for locality see Figure 21) B Gudeodiscus giardi giardi (Fischer, 1898) (for locality see Figure 19) C Gudeodiscus (Gudeodiscus) phlyarius (Mabille, 1887) (for locality see Figure 22) D Gudeodiscus (Gudeodiscus) fischeri (Gude, 1901) (for locality see Figure 17) E Gudeodiscus (Gudeodiscus) messageri raheemi Páll-Gergely & Hunyadi, ssp. n., 20080510A F–G Gudeodiscus (Gudeodiscus) villedaryi specimens collected at the same locality in two different dates: F November (2011/102) and G May (20090520A). All photos by B. Páll-Gergely.
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Barna Páll-Gergely, András Hunyadi, Jonathan Ablett, Hào Văn Lương, Fred Naggs, Takahiro Asami
Zookeys
Figure 29.Inner walls of male reproductive organs of Gudeodiscus and Halongella gen. n. species. A–B penis of Halongella schlumbergeri (Morlet, 1886), 20071122D C penis of Halongella fruhstorferi (Möllendorff, 1901) (for locality see Figure 25) D epiphallus of Gudeodiscus fischeri (Gude, 1901), (for locality see Figure 18) E epiphallus of Gudeodiscus giardi giardi (Fischer, 1898), (for locality see Figure 19) F penial caecum of Gudeodiscus (Gudeodiscus) messageri raheemi Páll-Gergely & Hunyadi, ssp. n. (for locality see Figure 20) G penial caecum of Gudeodiscus (Gudeodiscus) messageri raheemi ssp. n. (for locality see Figure 28E) H epiphallus of Halongella schlumbergeri (for locality see Figure 26) I Halongella fruhstorferi, (for locality see Figure 25) J spermatophore of Gudeodiscus (Gudeodiscus) fischeri, (for locality see Figure 17). All photos by B. Páll-Gergely.
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Barna Páll-Gergely, András Hunyadi, Jonathan Ablett, Hào Văn Lương, Fred Naggs, Takahiro Asami
Zookeys
Figure 30.Calcareous claws found in pockets on the inner penial wall of Gudeodiscus and Halongella gen. n. species. A–C, F Gudeodiscus (Gudeodiscus) villedaryi (Ancey, 1888) (for locality see Figure 23) D Gudeodiscus giardi giardi (Fischer, 1898) (for locality see Figure 19) E Gudeodiscus (Gudeodiscus) fischeri (Gude, 1901) (for locality see Figure 19) G–I Halongella schlumbergeri (Morlet, 1886) (for locality see Figure 29A–B). The claws in case of Gudeodiscus (Gudeodiscus) fischeri and Halongella schlumbergeri were too fragile for dissecting out, therefore drawings are presented. All images by B. Páll-Gergely.
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Barna Páll-Gergely, András Hunyadi, Jonathan Ablett, Hào Văn Lương, Fred Naggs, Takahiro Asami
Zookeys
Figure 7.Shells of Vietnamese Gudeodiscus and Halongella gen. n. species. A Gudeodiscus (Gudeodiscus?) francoisi (Fischer, 1898), MNHN 24601 (holotype of Plectopylis bavayi) B Gudeodiscus (Gudeodiscus?) francoisi, MNHN 9945 (holotype of Plectopylis francoisi) C Gudeodiscus (Gudeodiscus?) francoisi, NHMUK 1922.8.29.51 (holotype of Plectopylis lepida) D Halongella fruhstorferi (Möllendorff, 1901), SMF 9258 (lectotype) E Gudeodiscus (Gudeodiscus) giardi giardi (Fischer, 1898), MNHN IM-2010-12120 (syntype of Plectopylis congesta) F Gudeodiscus (Gudeodiscus) giardi giardi, NHMUK 1922.8.29.49 (syntype of Plectopylis congesta). Photos: T. Deli (A, B, E), H. Taylor (C, F) and E. Neubert (D). Scale represents 20 mm.
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Barna Páll-Gergely, András Hunyadi, Jonathan Ablett, Hào Văn Lương, Fred Naggs, Takahiro Asami
Zookeys
Figure 13.Parietal (A, C, E, F, H, J, L, N, P, R, T, V, X) and palatal (B, D, G, I, K, M, O, Q, S, U, W, Y) plicae and lamellae of Gudeodiscus species. A–B Gudeodiscus (Veludiscus) emigrans emigrans (Möllendorff, 1901), holotype (after Páll-Gergely and Hunyadi 2013) C–D Gudeodiscus (Veludiscus) emigrans quadrilamellatus Páll-Gergely, 2013 (after Páll-Gergely and Hunyadi 2013) E–K Gudeodiscus (Gudeodiscus?) francoisi (Fischer, 1898) E–F holotype of Plectopylis lepida Gude, 1901 (after Gude 1901b), G MNHN-IM-2012-2311 H–I holotype of Plectopylis bavayi Gude, 1901 (after Gude 1901a) J–K Plectopylis francoisi (after Gude 1899b) L–U Gudeodiscus (Gudeodiscus) giardi giardi (Fischer, 1898) L Plectopylis giardi (after Gude 1899a) M Plectopylis giardi (after Gude 1899b) N–O Plectopylis congesta Gude, 1899 (after Gude 1899a); P–Q Vn10-69 R–S Vn10-59 T–U 2011/85 V–Y Gudeodiscus (Gudeodiscus) villedaryi (Ancey, 1888) V–W holotype of Plectopylis (Endoplon) choanomphala Möllendorff, 1901 (after Gude 1901c) X–Y Vn10-47A. Inner views: D, F, I, K, M, O, Q, S, W, Y; Outer views: B, U.
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Barna Páll-Gergely, András Hunyadi, Jonathan Ablett, Hào Văn Lương, Fred Naggs, Takahiro Asami
Zookeys
Figure 39.Locations mentioned in literature on plectopylid taxonomy. 1 Muong-Bo 2 Phony-Tho 3 Trinh-Thuong 4 Muong-Hum 5 Nat-Son (Nhat Son) 6 Muong-Kong 7 Long-Ping 8 Pac-Kha 9 Ha-Giang 10 Tinh-Tuc 11 Cao-Bang 12 Déo-Ma-Phuc 13 Quang-Huyen 14 Ha-Lang 15 Cho-Ra 16 Bac-Khan 17 Nac-Ri 18 That-Khé 19 Cho-Moi 20 Lang-Son 21 Mansongebirge 22 Than-Moi 23 Bac-Ninh 24 Dong-Trieu 25 Bah-Mun 26 Kebao 27 Baie d’Along. The locations of “Col de Nuages” (Clouds Pass) could not be located. It is probably situated on Lao Kay Province, close to Muong-Hum.
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Barna Páll-Gergely, András Hunyadi, Jonathan Ablett, Hào Văn Lương, Fred Naggs, Takahiro Asami
Zookeys
Figure 42.Distribution of Gudeodiscus species. Legends: filled triangle: Gudeodiscus (Veludiscus) emigrans quadrilamellatus Páll-Gergely, 2013, empty circle: Gudeodiscus (Gudeodiscus) giardi giardi (Fischer, 1898), filled circle: Gudeodiscus (Gudeodiscus?) francoisi (Fischer, 1899), semi filled circle: co-occurrence of the latter two species.
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Barna Páll-Gergely, András Hunyadi, Jonathan Ablett, Hào Văn Lương, Fred Naggs, Takahiro Asami
Zookeys
Figure 7.Shells of Vietnamese Gudeodiscus and Halongella gen. n. species. A Gudeodiscus (Gudeodiscus?) francoisi (Fischer, 1898), MNHN 24601 (holotype of Plectopylis bavayi) B Gudeodiscus (Gudeodiscus?) francoisi, MNHN 9945 (holotype of Plectopylis francoisi) C Gudeodiscus (Gudeodiscus?) francoisi, NHMUK 1922.8.29.51 (holotype of Plectopylis lepida) D Halongella fruhstorferi (Möllendorff, 1901), SMF 9258 (lectotype) E Gudeodiscus (Gudeodiscus) giardi giardi (Fischer, 1898), MNHN IM-2010-12120 (syntype of Plectopylis congesta) F Gudeodiscus (Gudeodiscus) giardi giardi, NHMUK 1922.8.29.49 (syntype of Plectopylis congesta). Photos: T. Deli (A, B, E), H. Taylor (C, F) and E. Neubert (D). Scale represents 20 mm.
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Barna Páll-Gergely, András Hunyadi, Jonathan Ablett, Hào Văn Lương, Fred Naggs, Takahiro Asami
Zookeys
Figure 35.SEM images of radulae of Gudeodiscus species. A, D, G, J, M show the middle part of the radula B, E, H, K, N show the central tooth and the first 2–3 pairs of laterals C, F, I, L, O show the marginals. A–C Gudeodiscus (Gudeodiscus) giardi giardi (Fischer, 1898) (for locality see Figure 19) D–F Gudeodiscus (Gudeodiscus) messageri raheemi Páll-Gergely & Hunyadi, ssp. n. (for locality see Figure 20) G–I Gudeodiscus (Gudeodiscus) multispira (Möllendorff, 1883), China, Guangxi, Qingshan, Qingshan Zhen, Lipu Xian, ca 250 m, 24°26.189'N, 110°20.008'E, leg. Nakahara, Y., Ohara, K., Okubo, K. & Otani, J. U., 12.11.2004. J–L Gudeodiscus (Gudeodiscus) phlyarius (Mabille, 1887), Lạng Sơn Province, ca. km. 50 of road 1B, 10 km to Bình Gia, 21°53.911'N, 106°25.664'E, leg. Hemmen, Ch. & J., 01.04.2011. M–O Gudeodiscus (Gudeodiscus) villedaryi (Ancey, 1888), (for locality see Figure 23). All photos by B. Páll-Gergely.
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Barna Páll-Gergely, András Hunyadi, Jonathan Ablett, Hào Văn Lương, Fred Naggs, Takahiro Asami
Zookeys
Figure 45.Living specimens of Gudeodiscus (Gudeodiscus) giardi giardi (Fischer, 1898) (A), Cao Bằng Province, Hòa An District, Nguyễn Huệ Commune, small hill just outside of Khau Trang Village, 22°33.510'N, 106°10.294'E, leg. Naggs, F. et al. 22.06.2011.; and Halongella schlumbergeri (Morlet, 1886) (B), Halong Bay area, Vietnam. Photos: F. Naggs.
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Barna Páll-Gergely, András Hunyadi, Jonathan Ablett, Hào Văn Lương, Fred Naggs, Takahiro Asami
Zookeys
Figure 42.Distribution of Gudeodiscus species. Legends: filled triangle: Gudeodiscus (Veludiscus) emigrans quadrilamellatus Páll-Gergely, 2013, empty circle: Gudeodiscus (Gudeodiscus) giardi giardi (Fischer, 1898), filled circle: Gudeodiscus (Gudeodiscus?) francoisi (Fischer, 1899), semi filled circle: co-occurrence of the latter two species.
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Barna Páll-Gergely, András Hunyadi, Jonathan Ablett, Hào Văn Lương, Fred Naggs, Takahiro Asami
Zookeys
Figure 8.Shells of Vietnamese Gudeodiscus and Halongella gen. n. species. A Gudeodiscus (Gudeodiscus) giardi giardi (Fischer, 1898), MNHN 9946 (syntype of Plectopylis giardi) B Gudeodiscus (Gudeodiscus) villedaryi (Ancey, 1888), SMF 9279 (lectotype of Plectopylis choanomphala) C Gudeodiscus (Gudeodiscus) villedaryi, NHMUK 1930.9.12.38 (holotype of Plectopylis villedaryi) D Gudeodiscus (Gudeodiscus) villedaryi, Vn11-152, coll PGB E Gudeodiscus (Gudeodiscus) dautzenbergi (Gude, 1901), MNHN 24603 (holotype) F Gudeodiscus (Gudeodiscus) dautzenbergi, MNHN 24602 (holotype of Plectopylis persimilis). Photos: T. Deli (A, E, F), S. Hof (B), H. Taylor (C) and B. Páll-Gergely (D). Scale represents 20 mm.
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Barna Páll-Gergely, András Hunyadi, Jonathan Ablett, Hào Văn Lương, Fred Naggs, Takahiro Asami
Zookeys
Figure 9.Shells (A–B) and apertural views (C–R) of Vietnamese Gudeodiscus, Sicradiscus and Halongella gen. n. species. A Gudeodiscus (Gudeodiscus?) suprafilaris (Gude, 1908), MNHN 24586 (holotype?) B Gudeodiscus (Gudeodiscus?) suprafilaris, 2011/81 C Gudeodiscus (Gudeodiscus) phlyarius (Mabille, 1887), Vn11-156 D Gudeodiscus (Gudeodiscus) phlyarius (Mabille, 1887) (typical “anterides/gouldingi”), MNHN-IM-2012-2164 E Gudeodiscus (Gudeodiscus) messageri messageri (Gude, 1909), MNHN-IM-2012-2215 F Gudeodiscus (Gudeodiscus?) hemmeni Páll-Gergely & Hunyadi, sp. n., Vn10-103A G Gudeodiscus (Gudeodiscus?) anceyi (Gude, 1901), GS22 H Sicradiscus mansuyi (Gude, 1908), 20081116C I Gudeodiscus (Gudeodiscus) giardi giardi (Fischer, 1898), 2011/81 J Gudeodiscus (Gudeodiscus) villedaryi (Ancey, 1888), Vn11-151 K–L Gudeodiscus (Gudeodiscus) dautzenbergi (Gude, 1901), Vn10-44 M–N Halongella schlumbergeri (Morlet, 1886), MAA3 O Halongella fruhstorferi (Möllendorff, 1901), Vn11-171 P Gudeodiscus (Gudeodiscus) fischeri (Gude, 1901), Vn10-120 Q Gudeodiscus (Gudeodiscus) fischeri (Gude, 1901), 20090515C R Gudeodiscus (Gudeodiscus?) suprafilaris (Gude, 1908), 2011/81. All photos by B. Páll-Gergely except for Figure 9A (T. Deli). Scale represents 10 mm and refers to A and B.
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Barna Páll-Gergely, András Hunyadi, Jonathan Ablett, Hào Văn Lương, Fred Naggs, Takahiro Asami
Zookeys
Figure 13.Parietal (A, C, E, F, H, J, L, N, P, R, T, V, X) and palatal (B, D, G, I, K, M, O, Q, S, U, W, Y) plicae and lamellae of Gudeodiscus species. A–B Gudeodiscus (Veludiscus) emigrans emigrans (Möllendorff, 1901), holotype (after Páll-Gergely and Hunyadi 2013) C–D Gudeodiscus (Veludiscus) emigrans quadrilamellatus Páll-Gergely, 2013 (after Páll-Gergely and Hunyadi 2013) E–K Gudeodiscus (Gudeodiscus?) francoisi (Fischer, 1898) E–F holotype of Plectopylis lepida Gude, 1901 (after Gude 1901b), G MNHN-IM-2012-2311 H–I holotype of Plectopylis bavayi Gude, 1901 (after Gude 1901a) J–K Plectopylis francoisi (after Gude 1899b) L–U Gudeodiscus (Gudeodiscus) giardi giardi (Fischer, 1898) L Plectopylis giardi (after Gude 1899a) M Plectopylis giardi (after Gude 1899b) N–O Plectopylis congesta Gude, 1899 (after Gude 1899a); P–Q Vn10-69 R–S Vn10-59 T–U 2011/85 V–Y Gudeodiscus (Gudeodiscus) villedaryi (Ancey, 1888) V–W holotype of Plectopylis (Endoplon) choanomphala Möllendorff, 1901 (after Gude 1901c) X–Y Vn10-47A. Inner views: D, F, I, K, M, O, Q, S, W, Y; Outer views: B, U.
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Barna Páll-Gergely, András Hunyadi, Jonathan Ablett, Hào Văn Lương, Fred Naggs, Takahiro Asami
Zookeys
Figure 19.Reproductive anatomy of Gudeodiscus (Gudeodiscus) giardi giardi (Fischer, 1898). Locality information: Cao Bằng Province, Quảng Uyên N, 206–207 cross, 430 m, 22°42.737'N, 106°27.223'E, leg. Hunyadi, A., 16.11.2011. Scales represents 5 mm (left) and 2 mm (right).
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Barna Páll-Gergely, András Hunyadi, Jonathan Ablett, Hào Văn Lương, Fred Naggs, Takahiro Asami
Zookeys
Figure 28.Inner walls of the penis of Gudeodiscus Páll-Gergely, 2013 species. A Gudeodiscus (Gudeodiscus) phlyarius (Mabille, 1887) (typical fallax specimen, for locality see Figure 21) B Gudeodiscus giardi giardi (Fischer, 1898) (for locality see Figure 19) C Gudeodiscus (Gudeodiscus) phlyarius (Mabille, 1887) (for locality see Figure 22) D Gudeodiscus (Gudeodiscus) fischeri (Gude, 1901) (for locality see Figure 17) E Gudeodiscus (Gudeodiscus) messageri raheemi Páll-Gergely & Hunyadi, ssp. n., 20080510A F–G Gudeodiscus (Gudeodiscus) villedaryi specimens collected at the same locality in two different dates: F November (2011/102) and G May (20090520A). All photos by B. Páll-Gergely.
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Barna Páll-Gergely, András Hunyadi, Jonathan Ablett, Hào Văn Lương, Fred Naggs, Takahiro Asami
Zookeys
Figure 29.Inner walls of male reproductive organs of Gudeodiscus and Halongella gen. n. species. A–B penis of Halongella schlumbergeri (Morlet, 1886), 20071122D C penis of Halongella fruhstorferi (Möllendorff, 1901) (for locality see Figure 25) D epiphallus of Gudeodiscus fischeri (Gude, 1901), (for locality see Figure 18) E epiphallus of Gudeodiscus giardi giardi (Fischer, 1898), (for locality see Figure 19) F penial caecum of Gudeodiscus (Gudeodiscus) messageri raheemi Páll-Gergely & Hunyadi, ssp. n. (for locality see Figure 20) G penial caecum of Gudeodiscus (Gudeodiscus) messageri raheemi ssp. n. (for locality see Figure 28E) H epiphallus of Halongella schlumbergeri (for locality see Figure 26) I Halongella fruhstorferi, (for locality see Figure 25) J spermatophore of Gudeodiscus (Gudeodiscus) fischeri, (for locality see Figure 17). All photos by B. Páll-Gergely.
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Barna Páll-Gergely, András Hunyadi, Jonathan Ablett, Hào Văn Lương, Fred Naggs, Takahiro Asami
Zookeys
Figure 30.Calcareous claws found in pockets on the inner penial wall of Gudeodiscus and Halongella gen. n. species. A–C, F Gudeodiscus (Gudeodiscus) villedaryi (Ancey, 1888) (for locality see Figure 23) D Gudeodiscus giardi giardi (Fischer, 1898) (for locality see Figure 19) E Gudeodiscus (Gudeodiscus) fischeri (Gude, 1901) (for locality see Figure 19) G–I Halongella schlumbergeri (Morlet, 1886) (for locality see Figure 29A–B). The claws in case of Gudeodiscus (Gudeodiscus) fischeri and Halongella schlumbergeri were too fragile for dissecting out, therefore drawings are presented. All images by B. Páll-Gergely.