Gudeodiscus fischeri (Gude 1901)

Figure 2.Shells of Vietnamese Sicradiscus and Gudeodiscus species. A Sicradiscus mansuyi (Gude, 1908), NHMUK 1907.2.20.19 (syntype) B Gudeodiscus (Gudeodiscus?) anceyi (Gude, 1901), Tonkin, Bac-Kan, leg. Messager, MNHN 24600 (syntype) C Gudeodiscus (Gudeodiscus?) hemmeni Páll-Gergely & Hunyadi, sp. n., 2012/61, HNHM 97458 (holotype) D Gudeodiscus (Gudeodiscus?) hemmeni, Vn10-103 E Gudeodiscus (Gudeodiscus) fischeri (Gude, 1901), 20090519B, coll. PGB F Gudeodiscus (Gudeodiscus?) cyrtochilus (Gude, 1909), NHMUK 1922.8.29.59. (syntype). Photos: H. Taylor (A, F), T. Deli (B) and B. Páll-Gergely (C, D, E). Scale represents 10 mm.

Figure 31.Inner wall of the vagina of Sicradiscus and Gudeodiscus species. A Sicradiscus mansuyi (Gude, 1908), (for locality see Figure 27) B Gudeodiscus (Gudeodiscus) messageri raheemi Páll-Gergely & Hunyadi, ssp. n. (for locality see Figure 20) C Gudeodiscus (Gudeodiscus) phlyarius (Mabille, 1887), Vn11-157 D Gudeodiscus (Gudeodiscus) fischeri (Gude, 1901) (for locality see Figure 17).

Figure 34.SEM images of radulae of Gudeodiscus species. A, D, G, J, M show the middle part of the radula B, E, H, K, N show the central tooth and the first 2–3 pairs of laterals C, F, I, L, O show the marginals. A–C Gudeodiscus (Veludiscus) emigrans otanii Páll-Gergely & Hunyadi, 2013, China, Guangxi, Yizhou Shi, Aishan Xiang, Xiannuyan, ca 170 m, 24°29.292'N, 108°34.057'E, leg. Nakahara, Y., Ohara, K., Okubo, K. & Otani, J. U., 13.11.2004. D–F Gudeodiscus (Veludiscus) eroessi eroessi Páll-Gergely & Hunyadi, 2013, China, Guangxi, Guigang Shi, Guzhang Xiang, beyond Chuanshan village, ca 155 m, 23°20.848'N, 109°19.256'E, leg. Nakahara, Y., Ohara, K., Okubo, K. & Otani, J. U., 09.11.2004 G–I Gudeodiscus (Veludiscus) okuboi Páll-Gergely & Hunyadi, 2013, Guangxi, Guigang Shi, Guzhang Xiang, road to Wushan Xiang, ca 130 m, 23°21.178'N, 109°17.432'E, leg. Nakahara, Y., Ohara, K., Okubo, K. & Otani, J. U., 09.11.2004. J–L Gudeodiscus (Veludiscus) pulvinaris pulvinaris (Gould, 1859), China, Hong Kong Peak, leg. Miu Yeung, June 2013 M–O Gudeodiscus (Gudeodiscus) fischeri (Gude, 1901), (for locality see Figure 17). All photos by B. Páll-Gergely.

Figure 41.Distribution of Gudeodiscus species. Legends: filled circle: Gudeodiscus (Gudeodiscus?) suprafilaris (Gude, 1908), triangle: Gudeodiscus (Gudeodiscus) fischeri (Gude, 1901), empty square: Gudeodiscus (Gudeodiscus?) cyrtochilus (Gude, 1909).

Figure 39.Locations mentioned in literature on plectopylid taxonomy. 1 Muong-Bo 2 Phony-Tho 3 Trinh-Thuong 4 Muong-Hum 5 Nat-Son (Nhat Son) 6 Muong-Kong 7 Long-Ping 8 Pac-Kha 9 Ha-Giang 10 Tinh-Tuc 11 Cao-Bang 12 Déo-Ma-Phuc 13 Quang-Huyen 14 Ha-Lang 15 Cho-Ra 16 Bac-Khan 17 Nac-Ri 18 That-Khé 19 Cho-Moi 20 Lang-Son 21 Mansongebirge 22 Than-Moi 23 Bac-Ninh 24 Dong-Trieu 25 Bah-Mun 26 Kebao 27 Baie d’Along. The locations of “Col de Nuages” (Clouds Pass) could not be located. It is probably situated on Lao Kay Province, close to Muong-Hum.

Figure 3.Shells of Vietnamese Gudeodiscus species. A Gudeodiscus (Gudeodiscus) fischeri (Gude, 1901), Vn10-120, coll. PGB B Gudeodiscus (Gudeodiscus) fischeri, MNHN 24579 (holotype of Plectopylis fischeri) C Gudeodiscus (Gudeodiscus) fischeri, MNHN 24587 (holotype of Plectopylis tenuis) D Gudeodiscus (Gudeodiscus?) infralevis (Gude, 1908), MNHN 24604 (holotype of Plectopylis infralevis) E Gudeodiscus (Gudeodiscus?) infralevis, MNHN 24585 (holotype of Plectopylis soror). Photos: B. Páll-Gergely (A) and T. Deli (B–E). Scale represents 10 mm.

Figure 9.Shells (A–B) and apertural views (C–R) of Vietnamese Gudeodiscus, Sicradiscus and Halongella gen. n. species. A Gudeodiscus (Gudeodiscus?) suprafilaris (Gude, 1908), MNHN 24586 (holotype?) B Gudeodiscus (Gudeodiscus?) suprafilaris, 2011/81 C Gudeodiscus (Gudeodiscus) phlyarius (Mabille, 1887), Vn11-156 D Gudeodiscus (Gudeodiscus) phlyarius (Mabille, 1887) (typical “anterides/gouldingi”), MNHN-IM-2012-2164 E Gudeodiscus (Gudeodiscus) messageri messageri (Gude, 1909), MNHN-IM-2012-2215 F Gudeodiscus (Gudeodiscus?) hemmeni Páll-Gergely & Hunyadi, sp. n., Vn10-103A G Gudeodiscus (Gudeodiscus?) anceyi (Gude, 1901), GS22 H Sicradiscus mansuyi (Gude, 1908), 20081116C I Gudeodiscus (Gudeodiscus) giardi giardi (Fischer, 1898), 2011/81 J Gudeodiscus (Gudeodiscus) villedaryi (Ancey, 1888), Vn11-151 K–L Gudeodiscus (Gudeodiscus) dautzenbergi (Gude, 1901), Vn10-44 M–N Halongella schlumbergeri (Morlet, 1886), MAA3 O Halongella fruhstorferi (Möllendorff, 1901), Vn11-171 P Gudeodiscus (Gudeodiscus) fischeri (Gude, 1901), Vn10-120 Q Gudeodiscus (Gudeodiscus) fischeri (Gude, 1901), 20090515C R Gudeodiscus (Gudeodiscus?) suprafilaris (Gude, 1908), 2011/81. All photos by B. Páll-Gergely except for Figure 9A (T. Deli). Scale represents 10 mm and refers to A and B.

Figure 15.Parietal (A, C, E, F, H, J, L, N, O, Q) and palatal (B, D, G, I, K, M, P, R) plicae and lamellae of Gudeodiscus species. A–D Gudeodiscus (Gudeodiscus) infralevis (Gude, 1908) A–B holotype of Plectopylis infralevis (after Gude 1908) C–D Holotype of Plectopylis soror (after Gude 1908) E–G Gudeodiscus (Gudeodiscus?) cyrtochilus (Gude, 1909) E–F MNHN-IM-2012-2251 (two different specimens) G 2012/47 H–N Gudeodiscus (Gudeodiscus) fischeri (Gude, 1901) H–I Plectopylis fischeri (after Gude 1901a) J–K 20090515C L–M Vn10-120 N MNHN-IM-2012-2241 O–R Gudeodiscus (Gudeodiscus) fischeri (Gude, 1901) (identical with the holotype of tenuis) O–P after Gude (1901a) Q–R Vn10-28A (two different specimens). Inner views: B, D, I, M, P; Outer views: G, K, R.

Figure 17.Reproductive anatomy of Gudeodiscus (Gudeodiscus) fischeri (Gude, 1901). Locality information: Tuyên Quang Province, near Ton Hông, road #185 from Tuyên Quang to Vĩnh Lộc (formerly Chiêm Hóa) (NE of Tuyên Quang), leg. Hemmen, Ch. & J., 19.03.2011. Scale represents 5 mm.

Figure 18.Reproductive anatomy of Gudeodiscus (Gudeodiscus) fischeri (Gude, 1901) (typical tenuis specimen). Locality information: Bắc Kạn Province, Ba Bể Nat. Park, Hang Thẳm Kit 2 km from the look-out tower, 335 m, 22°24.686'N, 105°37.710'E, leg. Hunyadi, A., 19.11.2011. Scale represents 5 mm.

Figure 28.Inner walls of the penis of Gudeodiscus Páll-Gergely, 2013 species. A Gudeodiscus (Gudeodiscus) phlyarius (Mabille, 1887) (typical fallax specimen, for locality see Figure 21) B Gudeodiscus giardi giardi (Fischer, 1898) (for locality see Figure 19) C Gudeodiscus (Gudeodiscus) phlyarius (Mabille, 1887) (for locality see Figure 22) D Gudeodiscus (Gudeodiscus) fischeri (Gude, 1901) (for locality see Figure 17) E Gudeodiscus (Gudeodiscus) messageri raheemi Páll-Gergely & Hunyadi, ssp. n., 20080510A F–G Gudeodiscus (Gudeodiscus) villedaryi specimens collected at the same locality in two different dates: F November (2011/102) and G May (20090520A). All photos by B. Páll-Gergely.

Figure 29.Inner walls of male reproductive organs of Gudeodiscus and Halongella gen. n. species. A–B penis of Halongella schlumbergeri (Morlet, 1886), 20071122D C penis of Halongella fruhstorferi (Möllendorff, 1901) (for locality see Figure 25) D epiphallus of Gudeodiscus fischeri (Gude, 1901), (for locality see Figure 18) E epiphallus of Gudeodiscus giardi giardi (Fischer, 1898), (for locality see Figure 19) F penial caecum of Gudeodiscus (Gudeodiscus) messageri raheemi Páll-Gergely & Hunyadi, ssp. n. (for locality see Figure 20) G penial caecum of Gudeodiscus (Gudeodiscus) messageri raheemi ssp. n. (for locality see Figure 28E) H epiphallus of Halongella schlumbergeri (for locality see Figure 26) I Halongella fruhstorferi, (for locality see Figure 25) J spermatophore of Gudeodiscus (Gudeodiscus) fischeri, (for locality see Figure 17). All photos by B. Páll-Gergely.

Figure 30.Calcareous claws found in pockets on the inner penial wall of Gudeodiscus and Halongella gen. n. species. A–C, F Gudeodiscus (Gudeodiscus) villedaryi (Ancey, 1888) (for locality see Figure 23) D Gudeodiscus giardi giardi (Fischer, 1898) (for locality see Figure 19) E Gudeodiscus (Gudeodiscus) fischeri (Gude, 1901) (for locality see Figure 19) G–I Halongella schlumbergeri (Morlet, 1886) (for locality see Figure 29A–B). The claws in case of Gudeodiscus (Gudeodiscus) fischeri and Halongella schlumbergeri were too fragile for dissecting out, therefore drawings are presented. All images by B. Páll-Gergely.