Gudeodiscus messageri (Gude 1909)

Figure 5.Shells of Vietnamese Gudeodiscus species. A Gudeodiscus (Gudeodiscus) phlyarius (Mabille, 1887) (typical “fallax var. major”), MNHN-IM-2012-2155 B Gudeodiscus (Gudeodiscus) phlyarius, NHMUK 1922.8.29.55 (holotype of Plectopylis verecunda) C Gudeodiscus (Gudeodiscus) phlyarius, NHMUK 1922.8.29.58 (holotype of Plectopylis fallax) D Gudeodiscus (Gudeodiscus) messageri raheemi Páll-Gergely & Hunyadi, ssp. n., Vn10-76, coll. PGB E Gudeodiscus (Gudeodiscus) messageri raheemi ssp. n., NHMUK 20110370.1 (holotype) F Gudeodiscus (Gudeodiscus) messageri messageri (Gude, 1909), NHMUK 1922.8.29.53 (holotype) G Gudeodiscus (Gudeodiscus) messageri messageri NHMUK 1922.8.29.54 (syntype of Plectopylis messageri var. minor). Photos: B. Páll-Gergely (A, D), H. Taylor (B–C, E–G). Scale represents 10 mm.

Figure 9.Shells (A–B) and apertural views (C–R) of Vietnamese Gudeodiscus, Sicradiscus and Halongella gen. n. species. A Gudeodiscus (Gudeodiscus?) suprafilaris (Gude, 1908), MNHN 24586 (holotype?) B Gudeodiscus (Gudeodiscus?) suprafilaris, 2011/81 C Gudeodiscus (Gudeodiscus) phlyarius (Mabille, 1887), Vn11-156 D Gudeodiscus (Gudeodiscus) phlyarius (Mabille, 1887) (typical “anterides/gouldingi”), MNHN-IM-2012-2164 E Gudeodiscus (Gudeodiscus) messageri messageri (Gude, 1909), MNHN-IM-2012-2215 F Gudeodiscus (Gudeodiscus?) hemmeni Páll-Gergely & Hunyadi, sp. n., Vn10-103A G Gudeodiscus (Gudeodiscus?) anceyi (Gude, 1901), GS22 H Sicradiscus mansuyi (Gude, 1908), 20081116C I Gudeodiscus (Gudeodiscus) giardi giardi (Fischer, 1898), 2011/81 J Gudeodiscus (Gudeodiscus) villedaryi (Ancey, 1888), Vn11-151 K–L Gudeodiscus (Gudeodiscus) dautzenbergi (Gude, 1901), Vn10-44 M–N Halongella schlumbergeri (Morlet, 1886), MAA3 O Halongella fruhstorferi (Möllendorff, 1901), Vn11-171 P Gudeodiscus (Gudeodiscus) fischeri (Gude, 1901), Vn10-120 Q Gudeodiscus (Gudeodiscus) fischeri (Gude, 1901), 20090515C R Gudeodiscus (Gudeodiscus?) suprafilaris (Gude, 1908), 2011/81. All photos by B. Páll-Gergely except for Figure 9A (T. Deli). Scale represents 10 mm and refers to A and B.

Figure 12.Parietal (A, C, E, G–K, M, N, P–S, U, W) and palatal (B, D, F, L, P, T, V, X) plicae and lamellae of Gudeodiscus species. A–M Gudeodiscus (Gudeodiscus) phlyarius (Mabille, 1887) A–B Plectopylis phlyaria (after Gude 1901c) C–D Plectopylis moellendorffi (after Gude 1901c) E–F Vn10-49, G Vn09-24 H Vn10-56, I Vn9-16, spec.1 J Vn9-16, spec.2 K–M Plectopylis verecunda, MNHN 2012-2177 (3 different specimens) N–Q Gudeodiscus (Gudeodiscus) messageri messageri (Gude, 1909) N–O Plectopylis messageri (after Gude 1909) P MNHN-IM-2012-2162, Q MNHN-IM-2012-2165 R–V Gudeodiscus (Gudeodiscus) messageri raheemi ssp. n, R 20071116C, spec.1. S–T 20071116C, spec.2. U–V 20080509C W–X Vn10-104B. Inner views: B, D, F, O; Outer views: L, T, V, X.

Figure 43.Distribution of Gudeodiscus Páll-Gergely, 2013 species. Legends: filled triangle, top down: typical Gudeodiscus (Gudeodiscus) phlyarius (Mabille, 1887), filled triangle, top up: “Gudeodiscus phlyarius werneri Páll-Gergely, 2013” (synonym of phlyarius); empty triangle, top up: Gudeodiscus (Gudeodiscus) phlyarius populations showing transitional characters towards werneri in terms of shell shape; empty triangle, top down: atypical Gudeodiscus (Gudeodiscus) phlyarius; empty circle: Gudeodiscus messageri raheemi ssp. n., filled circle: Gudeodiscus (Gudeodiscus?) hemmeni sp. n. (in all localities it co-occurs with Gudeodiscus (Gudeodiscus) messageri raheemi ssp. n.); circle with filled triangle in the middle: co-occurrence of Gudeodiscus (Gudeodiscus) messageri raheemi ssp. n. and atypical Gudeodiscus (Gudeodiscus) phlyarius. The shaded area indicates the area inhabited by Gudeodiscus (Gudeodiscus) messageri messageri (Gude, 1909) and “anterides”, “fallax” and “gouldingi”-like populations of Gudeodiscus (Gudeodiscus) phlyarius. Filled square indicates the position of Phong-Tho, the type locality of Plectopylis verecunda Gude, 1909 (synonym of Gudeodiscus phlyarius). Numbers 1–3 refer to atypical populations assigned to Gudeodiscus (Gudeodiscus) phlyarius. For explanation, see text.

Figure 5.Shells of Vietnamese Gudeodiscus species. A Gudeodiscus (Gudeodiscus) phlyarius (Mabille, 1887) (typical “fallax var. major”), MNHN-IM-2012-2155 B Gudeodiscus (Gudeodiscus) phlyarius, NHMUK 1922.8.29.55 (holotype of Plectopylis verecunda) C Gudeodiscus (Gudeodiscus) phlyarius, NHMUK 1922.8.29.58 (holotype of Plectopylis fallax) D Gudeodiscus (Gudeodiscus) messageri raheemi Páll-Gergely & Hunyadi, ssp. n., Vn10-76, coll. PGB E Gudeodiscus (Gudeodiscus) messageri raheemi ssp. n., NHMUK 20110370.1 (holotype) F Gudeodiscus (Gudeodiscus) messageri messageri (Gude, 1909), NHMUK 1922.8.29.53 (holotype) G Gudeodiscus (Gudeodiscus) messageri messageri NHMUK 1922.8.29.54 (syntype of Plectopylis messageri var. minor). Photos: B. Páll-Gergely (A, D), H. Taylor (B–C, E–G). Scale represents 10 mm.

Figure 10.SEM images of Gudeodiscus shells. A protoconch of Gudeodiscus (Gudeodiscus) messageri raheemi Páll-Gergely & Hunyadi, ssp. n., Vn12-104, coll. HE B protoconch of Gudeodiscus (Gudeodiscus) villedaryi (Ancey, 1888), Vn11-163, coll. HE C–D sculpture of Gudeodiscus (Gudeodiscus) phlyarius (Mabille, 1887), Vn10-56, coll. HE E–F sculpture of Gudeodiscus (Gudeodiscus) phlyarius (Mabille, 1887) (typical fallax specimen), Vn11-187, coll HE. Images: B. Páll-Gergely.

Figure 12.Parietal (A, C, E, G–K, M, N, P–S, U, W) and palatal (B, D, F, L, P, T, V, X) plicae and lamellae of Gudeodiscus species. A–M Gudeodiscus (Gudeodiscus) phlyarius (Mabille, 1887) A–B Plectopylis phlyaria (after Gude 1901c) C–D Plectopylis moellendorffi (after Gude 1901c) E–F Vn10-49, G Vn09-24 H Vn10-56, I Vn9-16, spec.1 J Vn9-16, spec.2 K–M Plectopylis verecunda, MNHN 2012-2177 (3 different specimens) N–Q Gudeodiscus (Gudeodiscus) messageri messageri (Gude, 1909) N–O Plectopylis messageri (after Gude 1909) P MNHN-IM-2012-2162, Q MNHN-IM-2012-2165 R–V Gudeodiscus (Gudeodiscus) messageri raheemi ssp. n, R 20071116C, spec.1. S–T 20071116C, spec.2. U–V 20080509C W–X Vn10-104B. Inner views: B, D, F, O; Outer views: L, T, V, X.

Figure 20.Reproductive anatomy of Gudeodiscus (Gudeodiscus) messageri raheemi Páll-Gergely & Hunyadi, ssp. n. Locality information: Hòa Bình Province, ca. km 156 old road Hà Nội to Sơn La (right side off road), 20°46.000'N, 104°53.885'E, leg. Hemmen, Ch. & J., 15.10.2010. Scale represents 5 mm.

Figure 28.Inner walls of the penis of Gudeodiscus Páll-Gergely, 2013 species. A Gudeodiscus (Gudeodiscus) phlyarius (Mabille, 1887) (typical fallax specimen, for locality see Figure 21) B Gudeodiscus giardi giardi (Fischer, 1898) (for locality see Figure 19) C Gudeodiscus (Gudeodiscus) phlyarius (Mabille, 1887) (for locality see Figure 22) D Gudeodiscus (Gudeodiscus) fischeri (Gude, 1901) (for locality see Figure 17) E Gudeodiscus (Gudeodiscus) messageri raheemi Páll-Gergely & Hunyadi, ssp. n., 20080510A F–G Gudeodiscus (Gudeodiscus) villedaryi specimens collected at the same locality in two different dates: F November (2011/102) and G May (20090520A). All photos by B. Páll-Gergely.

Figure 29.Inner walls of male reproductive organs of Gudeodiscus and Halongella gen. n. species. A–B penis of Halongella schlumbergeri (Morlet, 1886), 20071122D C penis of Halongella fruhstorferi (Möllendorff, 1901) (for locality see Figure 25) D epiphallus of Gudeodiscus fischeri (Gude, 1901), (for locality see Figure 18) E epiphallus of Gudeodiscus giardi giardi (Fischer, 1898), (for locality see Figure 19) F penial caecum of Gudeodiscus (Gudeodiscus) messageri raheemi Páll-Gergely & Hunyadi, ssp. n. (for locality see Figure 20) G penial caecum of Gudeodiscus (Gudeodiscus) messageri raheemi ssp. n. (for locality see Figure 28E) H epiphallus of Halongella schlumbergeri (for locality see Figure 26) I Halongella fruhstorferi, (for locality see Figure 25) J spermatophore of Gudeodiscus (Gudeodiscus) fischeri, (for locality see Figure 17). All photos by B. Páll-Gergely.

Figure 31.Inner wall of the vagina of Sicradiscus and Gudeodiscus species. A Sicradiscus mansuyi (Gude, 1908), (for locality see Figure 27) B Gudeodiscus (Gudeodiscus) messageri raheemi Páll-Gergely & Hunyadi, ssp. n. (for locality see Figure 20) C Gudeodiscus (Gudeodiscus) phlyarius (Mabille, 1887), Vn11-157 D Gudeodiscus (Gudeodiscus) fischeri (Gude, 1901) (for locality see Figure 17).

Figure 35.SEM images of radulae of Gudeodiscus species. A, D, G, J, M show the middle part of the radula B, E, H, K, N show the central tooth and the first 2–3 pairs of laterals C, F, I, L, O show the marginals. A–C Gudeodiscus (Gudeodiscus) giardi giardi (Fischer, 1898) (for locality see Figure 19) D–F Gudeodiscus (Gudeodiscus) messageri raheemi Páll-Gergely & Hunyadi, ssp. n. (for locality see Figure 20) G–I Gudeodiscus (Gudeodiscus) multispira (Möllendorff, 1883), China, Guangxi, Qingshan, Qingshan Zhen, Lipu Xian, ca 250 m, 24°26.189'N, 110°20.008'E, leg. Nakahara, Y., Ohara, K., Okubo, K. & Otani, J. U., 12.11.2004. J–L Gudeodiscus (Gudeodiscus) phlyarius (Mabille, 1887), Lạng Sơn Province, ca. km. 50 of road 1B, 10 km to Bình Gia, 21°53.911'N, 106°25.664'E, leg. Hemmen, Ch. & J., 01.04.2011. M–O Gudeodiscus (Gudeodiscus) villedaryi (Ancey, 1888), (for locality see Figure 23). All photos by B. Páll-Gergely.

Figure 43.Distribution of Gudeodiscus Páll-Gergely, 2013 species. Legends: filled triangle, top down: typical Gudeodiscus (Gudeodiscus) phlyarius (Mabille, 1887), filled triangle, top up: “Gudeodiscus phlyarius werneri Páll-Gergely, 2013” (synonym of phlyarius); empty triangle, top up: Gudeodiscus (Gudeodiscus) phlyarius populations showing transitional characters towards werneri in terms of shell shape; empty triangle, top down: atypical Gudeodiscus (Gudeodiscus) phlyarius; empty circle: Gudeodiscus messageri raheemi ssp. n., filled circle: Gudeodiscus (Gudeodiscus?) hemmeni sp. n. (in all localities it co-occurs with Gudeodiscus (Gudeodiscus) messageri raheemi ssp. n.); circle with filled triangle in the middle: co-occurrence of Gudeodiscus (Gudeodiscus) messageri raheemi ssp. n. and atypical Gudeodiscus (Gudeodiscus) phlyarius. The shaded area indicates the area inhabited by Gudeodiscus (Gudeodiscus) messageri messageri (Gude, 1909) and “anterides”, “fallax” and “gouldingi”-like populations of Gudeodiscus (Gudeodiscus) phlyarius. Filled square indicates the position of Phong-Tho, the type locality of Plectopylis verecunda Gude, 1909 (synonym of Gudeodiscus phlyarius). Numbers 1–3 refer to atypical populations assigned to Gudeodiscus (Gudeodiscus) phlyarius. For explanation, see text.

Figure 24; A Perrottetia aquilonaria, specimen CUMZ 6278 B Perrottetia dugasti, syntype MNHN-IM-2000-30867 C Perrottetia megadentata, holotype CUMZ 6286 D Perrottetia unidentata, holotype CUMZ 6281 E Gudeodiscus messageri raheemi, holotype NHMUK 20110370.1 F Gudeodiscus sp., CUMZ collection.