Zospeum

Holotype from: Weigand AM, 2013. (Subterranean Biology 11: 45–53. doi: 10.3897/subtbiol.11.5966)Weigand AM (2013) New Zospeum species (Gastropoda, Ellobioidea, Carychiidae) from 980 m depth in the Lukina Jama–Trojama cave system (Velebit Mts., Croatia). Subterranean Biology 11: 45–53. doi: 10.3897/subtbiol.11.5966

Figure 3.Zospeum vasconicum sp. n., A–E different views of holotype (MNCN 15.05/60147H) F–I paratype shells (MNCN 15.05/60147P) H paratype shell with window cut in body whorl exposing columella. Scale bar 0.5 mm.

Summary[edit] Description: English: Columellar fold views of Zospeum tholussum. The columellar fold of the broken paratype specimen shell is shown in clockwise rotation. Date: 15 September 2013, 23:39:40. Source: Weigand A (2013). "New Zospeum species (Gastropoda, Ellobioidea, Carychiidae) from 980 m depth in the Lukina Jama – Trojama cave system (Velebit Mts., Croatia)". Subterranean Biology 11: 45--53. Pensoft Publishers. DOI:10.3897/subtbiol.11.5966. Author: Alexander M. Weigand.

Figure 2; Zospeum percostulatum sp. n. Different views of the holotype (A–C) and a paratype 1.58 mm height (D–G). Note the inner whitish/yellowish mass in the holotype corresponds to their soft parts and the different orientation of the longer aperture axis between A and D.

Summary[edit] Description: English: Zospeum biscaiense Gómez & Prieto, 1983. Distribution in Europe, scientific state of knowledge of 2012. Source description in English. Light colour indicated rare occurrences or regions where the species could eventually be expected to occur. Dark colour indicated relatively reliable occurrences, as well as regions where the species was expected to occur, and where the species was not known to be extremely rare. Dark colour indications were not necessarily based on published records. Species summary in AnimalBase Deutsch: Verbreitung in Europa, Kenntnisstand von 2012. Quellenbeschreibung auf deutsch Español: Distribución en Europa, nivel de conocimiento de 2012. Descripción de la fuente en español Français : Repartition en Europe, niveau de connaissance de 2012. Description de la source en français Italiano: Distribuzione in Europa, livello di conoscenza del 2012. Descrizione della fonte in italiano Português: Distribuição na Europa, nível de conhecimento de 2012. Descrição da fonte em português Čeština: Distribuce v Evropě, úroveň znalostí v roce 2012 Polski: Dystrybucja w Europie, poziom wiedzy z 2012 roku Русский: Распространение в Европе, уровень знаний в 2012 году. Date: 28 September 2012. Source: Distributional range map as published in Welter-Schultes, F. 2012: European non-marine molluscs, a guide for species identification. Pp A1-A3, 1-679, Q1-Q78. Author: Francisco Welter-Schultes.

from: Weigand AM, 2013. (Subterranean Biology 11: 45–53. doi: 10.3897/subtbiol.11.5966)Weigand AM (2013) New Zospeum species (Gastropoda, Ellobioidea, Carychiidae) from 980 m depth in the Lukina Jama–Trojama cave system (Velebit Mts., Croatia). Subterranean Biology 11: 45–53. doi: 10.3897/subtbiol.11.5966

Figure 4.Zospeum vasconicum sp. n., A, F–I shells from Cueva Silibranka-2 in frontal view (RMNH.5003915) B–E different views of specimen A; H shell with window cut in body whorl exposing columella. Scale bar 0.5 mm.

The Lukina Jama–Trojama cave system. Overview of the geographical position and 3D cave cross-section. In the latter, the region of collected shells (1) and the collection site of the living specimen of Zospeum tholussum (2) are indicated. The 3D cross-section was provided by D. Bakšić et al. (2010), Croatian Speleological Server, http://www.speleologija.hr/lukinajama . Photos were taken by J. Bedek.

Summary[edit] Description: English: Paratype of Zospeum percostulatum. Date: 4 December 2017, 20:33:34. Source: A morphological gap for Iberian Zospeum filled: Zospeum percostulatum sp. n. (Gastropoda, Eupulmonata, Carychiidae) a new species from Asturias (Spain). Author: Alvaro Alonso, Carlos E. Prieto, Sergio Quiñonero-Salgado, Emilio Rolán.

Figure 5.Zospeum vasconicum sp. n., Shells from Cueva de Otxas (A–F RMNH.5003916); and Cueva del Cranéo (G–I UPV/EHU-FC: 556). B–C different views of A; F, H shells with window cut in body whorl exposing columella. Scale bar 0.5 mm.

Summary[edit] Description: English: Holotype and paratypes of Zospeum tholussum. The holotype (former living specimen) is marked with a solid line; five paratype specimens (shells) are surrounded by dotted lines. Date: 15 September 2013, 23:39:40. Source: Weigand A (2013). "New Zospeum species (Gastropoda, Ellobioidea, Carychiidae) from 980 m depth in the Lukina Jama – Trojama cave system (Velebit Mts., Croatia)". Subterranean Biology 11: 45--53. Pensoft Publishers. DOI:10.3897/subtbiol.11.5966. Author: Alexander M. Weigand.

Figure 4; Zospeum percostulatum sp. n. Frontal views of three paratype shells (A–C 1.41, 1.80, 1.58 mm) and a shell from Cueva Collubina (D 1.49 mm) [ZUPV]. Note the robust, smooth columella through the window opened in the body whorl (C) and closer, smaller riblets in D.

Figure 2.Map indicating geographic position of massifs and caves with Zospeum species. Dot in Triangle, Zospeum zaldivarae sp. n.: Cueva de Las Paúles (locus typicus); Dot in circle, Zospeum vasconicum sp. n.: Cueva de la Ermita de Sandaili (locus typicus); Simple orange circles, Zospeum vasconicum sp. n. localities: Cueva de Otxas; Cueva Silibranka-2; Cueva del Cranéo.

Summary[edit] Description: English: Holotype and paratypes of Zospeum tholussum. The holotype (former living specimen) is marked with a solid line; five paratype specimens (shells) are surrounded by dotted lines. Date: 15 September 2013, 23:39:40. Source: Weigand A (2013). "New Zospeum species (Gastropoda, Ellobioidea, Carychiidae) from 980 m depth in the Lukina Jama – Trojama cave system (Velebit Mts., Croatia)". Subterranean Biology 11: 45--53. Pensoft Publishers. DOI:10.3897/subtbiol.11.5966. Author: Alexander M. Weigand. Other versions: : This file has been extracted from another file: Holotype and paratypes of Zospeum tholussum - SubtBiol-011-045-g002.jpg : .

Figure 3; Zospeum percostulatum sp. n. Frontal view of a paratype (A, 1.42 mm height) [MHNS] and some partial views (B–G): B protoconch C microsculpture of the protoconch D detail selected in C E suture and costulae of the last whorl F shape of costulae G microsculpture of the teleoconch.

Figure 1.Measurement on Zospeum shells in frontal view. Abbreviations: HLW height of last whorl PH peristome height PD peristome diameter SA spire angle SD shell diameter SH shell height.

Summary[edit] Description: English: A live individual of Zospeum tholussum is seen in its natural habitat. From the paper "New Zospeum species (Gastropoda, Ellobioidea, Carychiidae) from 980 m depth in the Lukina Jama – Trojama cave system (Velebit Mts., Croatia)". Date: 2013. Source: Weigand A (2013) New Zospeum species (Gastropoda, Ellobioidea, Carychiidae) from 980 m depth in the Lukina Jama–Trojama cave system (Velebit Mts., Croatia). Subterranean Biology 11: 45-53. doi:10.3897/subtbiol.11.5966. Author: J. Bedek, Alexander M. Weigand. Permission(Reusing this file): https://www.pensoft.net/about.php (Creative Commons - attribution- 3.0). This image appeared on English Wikipedia's Main Page in the Did you know? column on 25 October 2013 (see archives)..

Figure 8.Habitat of Zospeum species. A–B Moist muddy layer with Zospeum vasconicum sp. n. in Cueva Arrikrutz; Prov. Gipuzkoa, Natural Park of Aizkorri-Araotz, Oñate C Muddy sediment matrix of Zospeum zaldivarae sp. n. habitat in Cueva de Las Paúles (locus typicus) with congener (Zospeum suarezi) in view.

Summary[edit] Description: English: Holotype of Zospeum tholussum. Cropped from Fig. 2, which is also available as File:Holotype and paratypes of Zospeum tholussum - SubtBiol-011-045-g002.jpg. Date: 15 September 2013, 23:39:40. Source: Weigand A (2013). "New Zospeum species (Gastropoda, Ellobioidea, Carychiidae) from 980 m depth in the Lukina Jama – Trojama cave system (Velebit Mts., Croatia)". Subterranean Biology 11: 45--53. Pensoft Publishers. DOI:10.3897/subtbiol.11.5966. Author: Alexander M. Weigand.

Figure 6.Zospeum zaldivarae sp. n., A, E–F different views of holotype shell (MNCN 15.05/60148H) D aperture in slightly oblique view showing apertural barriers.

Summary[edit] Description: English: Holotype and paratypes of Zospeum tholussum. The holotype (former living specimen) is marked with a solid line; five paratype specimens (shells) are surrounded by dotted lines. Date: 15 September 2013, 23:39:40. Source: Weigand A (2013). "New Zospeum species (Gastropoda, Ellobioidea, Carychiidae) from 980 m depth in the Lukina Jama – Trojama cave system (Velebit Mts., Croatia)". Subterranean Biology 11: 45--53. Pensoft Publishers. DOI:10.3897/subtbiol.11.5966. Author: Alexander M. Weigand. Other versions: This file has an extracted image: File:Holotype of Zospeum tholussum - SubtBiol-011-045-g002 (cropped).jpg..

Figure 7.Zospeum zaldivarae sp. n., A–F different views of edentate paratype shell (RMNH.234152) D aperture in oblique view G paratype shell (UPV/EHU-FC: 72) with window in body whorl exposing columella.

Figure 8.Habitat of Zospeum species. A–B Moist muddy layer with Zospeum vasconicum sp. n. in Cueva Arrikrutz; Prov. Gipuzkoa, Natural Park of Aizkorri-Araotz, Oñate C Muddy sediment matrix of Zospeum zaldivarae sp. n. habitat in Cueva de Las Paúles (locus typicus) with congener (Zospeum suarezi) in view.

Figure 6; Zospeum cf. vasconicum Prieto, De Winter, Weigand, Gómez & Jochum, 2015. Ex. Z. suarezi paratype, NHMW-MO 75000-E-48815, Cueva Hernialde, Guipuzcoa, assessed in Gittenberger (1980) („NWW-Coll. Edlauer 48815, ex. Coll. Robić/1shell”).